Mar 31, 2026 filings. Quarterly 13F reports arrive up to 45 days after quarter end; this register is filed through May 29, 2026.
195 institutional managers reported holding APPN for the quarter ended Mar 31, 2026. Together they hold about 53% of shares outstanding; the top 10 hold 34%. 31 opened new positions; 79 closed out. Complete ranked register from SEC 13F filings.
| manager | shares | % out | value (Mar 31, 2026 close) | change | held since |
|---|---|---|---|---|---|
| Vanguard Group | 6.18M | 8.37% | $148.8M | added +1.3% | 2017 Q2 |
| RPD Fund Management LLC (focused) | 3.92M | 5.31% | $94.4M | opened | 2026 Q1 |
| Fivespan Partners, LP (focused) | 3.51M | 4.76% | $84.6M | added +155.3% | 2025 Q4 |
| BlackRock | 3.23M | 4.37% | $77.7M | trimmed -1.5% | 2017 Q2 |
| Lead Edge Capital Management, LLC (focused) | 1.91M | 2.58% | $45.9M | added +18.5% | 2024 Q2 |
| Morgan Stanley | 1.55M | 2.09% | $37.2M | added +8.9% | 2017 Q2 |
| Arrowstreet Capital | 1.52M | 2.06% | $36.6M | added +16.5% | 2024 Q4 |
| First Trust Advisors LP | 1.36M | 1.84% | $32.7M | added +16.9% | 2018 Q3 |
| State Street | 1.02M | 1.39% | $24.7M | added +4.4% | 2017 Q3 |
| Geode Capital | 985.5k | 1.33% | $23.7M | unchanged | 2017 Q3 |
| AQR Capital | 957.6k | 1.30% | $23.1M | added +42.5% | 2023 Q4 |
| Baillie Gifford | 905.9k | 1.23% | $21.8M | trimmed -3.7% | 2019 Q1 |
| Mudita Advisors LLP (focused) | 839.4k | 1.14% | $20.2M | opened | 2026 Q1 |
| Jacobs Levy Equity Management, INC | 603.4k | 0.82% | $14.5M | trimmed -2.9% | 2025 Q1 |
| Acadian Asset Management | 520.4k | 0.70% | $12.5M | trimmed -15.1% | 2024 Q3 |
| Citadel Advisors | 518.0k | 0.70% | $12.5M | trimmed -6.3% | 2024 Q4 |
| D. E. Shaw | 447.8k | 0.61% | $10.8M | trimmed -59.7% | 2018 Q2 |
| Nuveen | 415.0k | 0.56% | $10.0M | trimmed -40.2% | 2019 Q2 |
| Squarepoint Ops LLC | 390.9k | 0.53% | $9.4M | added +32.1% | 2021 Q4 |
| Prelude Capital Management, LLC | 390.3k | 0.53% | $9.4M | opened | 2025 Q2 |
| FMR (Fidelity) | 380.7k | 0.52% | $9.2M | added +18% | 2019 Q1 |
| Charles Schwab Investment Management | 370.7k | 0.50% | $8.9M | added +5.6% | 2018 Q3 |
| UBS | 370.3k | 0.50% | $8.9M | added +109.2% | 2017 Q3 |
| Mirae Asset Global Etfs Holdings LTD. | 358.8k | 0.49% | $8.6M | trimmed -34.1% | 2021 Q1 |
| Northern Trust | 344.3k | 0.47% | $8.3M | added +2.1% | 2017 Q3 |
| Trexquant Investment LP | 305.9k | 0.41% | $7.4M | added +97% | 2020 Q4 |
| National Philanthropic Trust | 299.5k | 0.41% | $7.2M | unchanged | 2024 Q1 |
| Qube Research & Technologies LTD | 292.7k | 0.40% | $7.1M | trimmed -38.9% | 2024 Q4 |
| Goldman Sachs | 285.3k | 0.39% | $6.9M | trimmed -30.5% | 2018 Q1 |
| Connor, Clark & Lunn Investment Management LTD. | 208.6k | 0.28% | $5.0M | trimmed -43.2% | 2024 Q1 |
| Jane Street | 194.5k | 0.26% | $4.7M | trimmed -22.7% | 2020 Q2 |
| HRT Financial LP | 190.0k | 0.26% | $4.6M | added +78.9% | 2025 Q3 |
| Algert Global LLC | 179.6k | 0.24% | $4.3M | added +22.9% | 2023 Q1 |
| CPP Investment Board | 179.5k | 0.24% | $4.3M | trimmed -8.7% | 2024 Q2 |
| Bank OF America CORP /DE/ | 176.6k | 0.24% | $4.3M | added +8% | 2017 Q4 |
| Tejara Capital LTD | 173.3k | 0.23% | $4.2M | opened | 2026 Q1 |
| Harber Asset Management LLC (focused) | 147.4k | 0.20% | $3.6M | trimmed -22.2% | 2023 Q4 |
| Point72 | 147.2k | 0.20% | $3.5M | trimmed -36.1% | 2019 Q3 |
| ExodusPoint | 146.0k | 0.20% | $3.5M | opened | 2026 Q1 |
| Engineers Gate Manager LP | 131.0k | 0.18% | $3.2M | trimmed -14.8% | 2024 Q1 |
| BNP Paribas Financial Markets | 128.1k | 0.17% | $3.1M | trimmed -48.2% | 2018 Q2 |
| Russell Investments Group, LTD. | 123.1k | 0.17% | $3.0M | trimmed -49.4% | 2023 Q4 |
| BNY Mellon | 120.4k | 0.16% | $2.9M | trimmed -2.2% | 2018 Q1 |
| Two Sigma | 119.1k | 0.16% | $2.9M | trimmed -62.7% | 2021 Q1 |
| Gotham Asset Management | 95.6k | 0.13% | $2.3M | added +98.5% | 2025 Q2 |
| SEI Investments CO | 85.1k | 0.12% | $2.1M | added +7.7% | 2025 Q2 |
| LOS Angeles Capital Management LLC | 80.6k | 0.11% | $1.9M | added +106.7% | 2025 Q3 |
| Swiss National Bank | 76.3k | 0.10% | $1.8M | unchanged | 2019 Q2 |
| Boothbay Fund Management, LLC | 75.2k | 0.10% | $1.8M | opened | 2026 Q1 |
| Barclays PLC | 74.7k | 0.10% | $1.8M | trimmed -43.6% | 2017 Q2 |
Largest 50 of 195 shown; the full ranked, searchable register is at boothcheck.com/holders/APPN. Related filers are grouped using the filings' own included-manager declarations. Historically, several managers opening new positions in the same quarter said almost nothing about the quarter that followed (59,118 cases since 2013). A filed record, not a forecast. Nothing here is investment advice.